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Sakura is the princess of Clow kingdom, which is ruled by her older brother, King Tōya. Her childhood friend Syaoran is a young archaeologist. Sakura is revealed to have strange powers when she has a vision of a mysterious symbol and places she has never seen. Meanwhile, Syaoran discovers the same symbol at the ruins he is excavating. He sees Sakura standing on the symbol on the ground. Ghostly wings appear on her back and a mysterious force begins to pull her into the walls of the ruins. Syaoran rescues her in time, but her wings are scattered across dimensions. The High Priest of Clow Kingdom, Yukito, immediately realizes that Sakura's "wings" were the manifestation of her soul and memories; without them, she will die. In order to save Sakura, Syaoran must journey to retrieve her wing's feathers, the fragments of her memories.

Yukito sends Syaoran and the unconscious Sakura to the Dimensional Witch, Yūko, who is one of the main characters in xxxHolic. There he meets Kurogane, a rough-mannered ninja banished from his world by Princess Tomoyo, and Fye D. Flourite, a magician who fled his world to avoid King Ashura. Each of them must pay with what he values most in order to gain the power to cross dimensions. For Kurogane, it is his sword Ginryū, and for Fye, it is the tattoo on his back which regulates his control of his magic. Syaoran, on the other hand, must pay with his relationship with Sakura: even if he is able to retrieve all of her memories, she will never remember anything about him or their relationship. (This sacrifice also pays Sakura's "toll" to Yūko, because what Sakura values most are her memories of and with Syaoran.) Only when the three agree to her terms does Yūko present them with the power to cross dimensions; a white creature named Mokona Modoki.

While traversing through worlds to find Sakura's feathers, the four travellers and Mokona are forced to overcome many dangers and opponents, some of whom are figures from Syaoran's past who wish to collect Sakura's feathers for their own reasons as they are seen to be objects that hold great power.

The manga adopts darker and more complex tones in the latter half of the story.

Because of the crossover characters with xxxHolic, the two manga occasionally intertwine with each other.

The hero of the story. He (at the beginning) does not have any magical power, but is an excellent martial artist and an aspiring archaeologist with extensive knowledge of various languages, histories, and artifacts. In Tsubasa: Reservoir Chronicle Syaoran is blind in his right eye. In the early stages of his life he is emotionally distressed, probably due to his true self being locked away while controlling him unawarely, but that later changes. His heart, which was given to him by the real Syaoran, has now been returned to its owner. He now only acts to fulfill Fei Wong Reed's plans to gather all of Sakura's feathers. A crossover character from Cardcaptor Sakura.

Syaoran is an archaeologist who is excavating ruins in the country of Clow and the adopted son of Fujitaka. In the anime, he is voiced by Miyu Irino in Japanese, and in English by Jason Liebrecht. Other than having a completely different family line, Syaoran is very mature and polite, unlike his more arrogant incarnation in Cardcaptor Sakura. However, he still maintains the overtly serious streak, which often makes him believe in almost anything, making him come off as a greatly gullible person.

In the series, Syaoran is shown to be an excellent fighter, centered around his devastating kicks (taught to him by Seishirou Sakurazuka. This is to compensate for his lack of reliable depth perception due to being blind in his right eye. He was later trained to sense attacks coming from the left by Kurogane). He is also an experienced archaeologist, with the attendant skills and experience of a hardened adventurer somewhat reminiscent of Indiana Jones. In Outo, he receives a sword named Hien ("scarlet flame"), whose blade can be made to burst into flame, and is trained in swordsmanship by Kurogane. In the Tsubasa Character Guide, Syaoran's fighting abilities are given a score of 380/500, only 20 points behind Fai and making him the third-strongest fighter at the start of the series.

The Princess of Clow Country. Sakura is a cheerful, selfless, and strong-hearted young woman. She states if there was ever something that she could do, she would try her absolute best. Her wings are the manifestation of her soul, namely her heart and memories. She possesses a mysterious power, manifest in the magical powers of the scattered feathers, which is sought by Fei Wong Reed. Much later on in the story, something shocking is found out about her. A crossover character from Cardcaptor Sakura.

Sakura as seen in Tsubasa ChronicleAt the start of the series in both manga and anime, Sakura is the princess of the Kingdom of Clow who has feelings for Syaoran, her childhood friend.

When she is about to confess her feelings, a mysterious force causes her own dormant magical powers to manifest in the form of feathered wings. When her feathers are scattered across time and space, Syaoran vows to recover all of them because each feather represents a memory Sakura has had since birth and are connected to her heart; without the feathers, she will die.

Fei-Wong Reed (the mysterious villain in the series) reflects on the ability of people like the High Priest Yukito, Princess Tomoyo of Nihon, and Yūko to cross dimensions and comments that the power concealed within Sakura and the ruins in Clow Country surpasses them all.

In Tsubasa: Tokyo Revelations, Sakura Kinomoto herself makes a cameo appearance.

A relationship between Sakura, Kimihiro Watanuki (to whom she shares a birthday) and Kohane Tsuyuri has been hinted at, in a similar way to Syaoran's relationship to Watanuki.

Piczo